OK;
Velocity is a function of average pressure over time. Imagine your plenum is the same size as your barrel. If your pressure in your plenum is 2 BAR then it contains twice as much air as your barrel. If your pressure is 4 BAR then it contains four times as much air as your barrel. Now imagine your plenum is twice the volume of your barrel. If your pressure is 1 BAR it contains twice as much air as your barrel. If your pressure is 2 BAR then it contains 4 times as much air as your barrel. Simple enough.
Pressure is stored energy. More pressure, more energy. When the pressure leaves a vessel the vessel cools off. This is energy leaving the vessel as heat. One liter of air stored at 2 BAR contains the same amount of energy as two liters of air stored at one BAR. This is true in a vacuum. It is also true when there is no differential in pressure because the energy being considered is actually heat.
To keep things simple, imagine the following is taking place in a vacuum. Imagine a barrel that is plugged at the far end with a pellet. Suppose you put the air into that barrel from a plenum that was 1/4 times as big as the barrel and was pressurized to 4 BAR. The pressure at the moment the pellet leaves the barrel is 1 BAR. This is also the average pressure in the barrel over the time the pellet traveled down it. Now imagine the same situation but the plenum is 1/2 the volume of the barrel and the pressure is only 2 BAR. At the moment the pellet leaves the barrel the pressure is still 1 BAR more importantly the average pressure in the barrel over the time the pellet traveled down it is also 1 BAR. Larger container, lower pressure, same velocity.
You can scale this to a plenum which is any size with any initial pressure. Imagine a plenum that is the same size as the barrel and is pressurized to 2 BAR. At the time the pellet exits the barrel the pressure is reduced to 1 BAR because the volume of the whole system (plenum plus barrel) has doubled. Suppose you increase that plenum to twice the size of the barrel and keep the pressure the same. At the time the pellet exits the pressure is 4 BAR/3 (33% higher). The average pressure over the time in the barrel is 33% greater. We could reduce the pressure and wind up with a velocity equal to the first example in this paragraph. With the same plenum (twice as big as the barrel) we can reduce the pressure to 3 BAR and end up with 1 BAR of average pressure at the time the pellet exits the barrel (3/3=1).
So what matters is average pressure over time and that number is always the same as the muzzle pressure.
I'm sorry I tried to make this easy to digest but it is hard to talk about, without doing calculus.