This is like whack-a-mole.
I cleaned up the regulator and changed the three seals that Rodeo mentioned. Like Rodeo's, the o-ring on the piston was dry, compressed and brittle. Reassembled it all and it leaked from the front of "part 38'" as XTX calls it.
Stripped it again and replaced the two o-rings on the front of part 38, reassembled it and now it leaks from the back of part 38.
I guess tomorrow, I'll strip it again and replace whatever seal or seals there are in the back end of part 38.
This has been pretty stupid. So far the fill port leaked - fixed it, the bottle seal leaked, fixed it, the regulator leaked, fixed it, front seals on part38 leaked, fixed it.
And it still leaks!